Papers / Books
From the Columbia University Press

The Adelie Penguin: Bellwether of Climate Change

Dr. David G. Ainley, author

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The Adélie penguin is one of the best-studied birds in the world and is the subject of research programs from a dozen nations interested in monitoring changes in the environment and the food webs of the Southern Ocean. This species’ population has been changing dramatically over the past few decades coincident with a general warming of the maritime portion of Antarctica. When the sea-ice is seen to decline so does the population of Adélie penguins. Further south, however, the population is increasing.

This book summarizes our present ecological knowledge of this polar seabird. In so doing, David Ainley describes the ecological factors important to its life history and details the mechanisms by which it is responding to climate change. The author also chronicles the history of research on Adélie penguins, beginning with the heroic expeditions at the beginning of the twentieth century.

From Tumblehome

Waiting for Joey

Jean Pennycook, author

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Join a teacher-scientist as she observes Antarctic penguins over ten breeding seasons.

How do Adelie penguins live and adapt to the changing climatic conditions of Antarctica? The author relates 10 years of observation, tracing the story of one penguin she calls Joey. Children will relate to both the penguins and the author’s sense of adventure as she becomes an expert penguin scientist. Richly illustrated with photos taken on-site at the research station in Antarctica.


If you would like to have a copy, please contact David Ainley 

Natural history Patterns
Ainley, D.G. 2007. A letter from Cape Royds: of penguins and men. Pp210-226 in Antarctica: Life on the Ice (S.F. Rogers, Ed.). Traveler’s Tales Books, Sola House, Palo Alto CA. view PDF

Ainley, D.G., S.S. Jacobs, C.A. Ribic & I. Gaffney. 1998. Seabird distribution and oceanic features of the Amundsen and southern Bellingshausen seas.  Antarctic Science 10: 111-123 . view PDF

Ainley, D.G., N. Nur & E. J. Woehler. 1995.  Factors affecting the size and distribution of pygoscelid penguin colonies in the Antarctic.  Auk 112: 171-182 . view PDF

Ainley, D.G., C.T. Tynan & I. Stirling. 2003. Sea ice: a critical habitat for polar marine mammals and birds. Pp 240-266inSea Ice: An Introduction to its Physics, Biology, Chemistry and Geology (D.N. Thomas & G.S. Dieckmann, Eds.). Blackwell Science, London.

Ainley, D.G., E.J. Woehler & A. Lescroël. 2017. Birds and Antarctic sea ice. In: Sea Ice (3rd Edition) (D.N. Thomas, Ed.). John Wiley & Sons, Ltd., London.

Ballard, G., D.G. Ainley, C.A. Ribic & K.R. Barton. 2001. Effect of instrument attachment on foraging trip duration and nesting success of Adélie penguins. Condor 103: 481-490. view PDF

Bost, C.A., C. Cotté, F. Bailleul, Y. Cherel, J.B. Charrassin, C. Guinet, D.G. Ainley,  H. Weimerskirch. 2009. The importance of oceanographic fronts to marine birds and mammals of the southern oceans. Journal of Marine Systems 78: 363–376. view PDF

Dugger, K.M., D.G. Ainley, P. O’B. Lyver, K.M. Barton, G. Ballard. 2010. Survival differences and the effect of environmental instability on breeding dispersal in an Adélie penguin meta-population. Proc Natl Acad Sci, doi 10.1073 pnas 1000623107. view PDF

Dugger, K.M., G. Ballard, Ainley, D.G. & K.J. Barton. 2006.  Flipper-band effects on the foraging behavior and survival of Adélie Penguins. Auk 123: 858–869. view PDF

LaRue, M.A., H.J. Lynch, P.O’B Lyver, K. Barton, D.G Ainley, A. Pollard, W.R. Fraser & G. Ballard. 2014. A method for estimating population colony sizes of Adélie penguins using remote sensing imagery. Polar Biology, doi 10.1007/s00300-014-1451-8 view PDF

Lescroël, A., K.M. Dugger, G. Ballard & Ainley, D.G. 2009. Effects of individual quality, reproductive success and environmental variability on survival of a long-lived seabird. Journal of Animal Ecology 78(4): 798-806. view PDF

Lyver POB, Barron M, Barton KJ, Ainley DG, Pollard A, Gordon S, McNeill S, Ballard G & Wilson P.R. 2014. Trends in the breeding population of Adélie penguins in the Ross Sea, 1981–2012: a coincidence of climate and resource extraction effects. PLoS ONE, 9(3): e91188. doi:10.1371/journal.pone.00911 view PDF

Petersen, S.L., G.M. Branch, D.G. Ainley,  P. D. Boersma, J. Cooper & E.J. Woehler. 2005. Is flipper banding of penguins a problem? Marine Ornithology 33: 75–79. view PDF

Ribic, C.A., D.G. Ainley,  R.G. Ford, W.R. Fraser, C.T. Tynan & E.J. Woehler. 2011. Water masses, ocean fronts, and the structure of Antarctic seabird communities: putting the eastern Bellingshausen Sea in perspective. Deep-Sea Research II 58: 1695-1709, view PDF

Russell, D.G.D., Sladen, W.J.L. and Ainley, D.G. 2012. Dr. George Murray Levick (1876-1956): unpublished notes on the sexual habits of the Adélie penguin. Polar Record, Available on CJO doi:10.1017/S0032247412000216. view PDF

Spear, L.B. & Ainley, D.G. 1998. Morphological differences relative to ecological segregation in petrels (Family: Procellariidae) of the Southern Ocean and tropical Pacific. Auk 115: 1017-1033. view PDF

Tynan, C.T., Ainley, D.G. & I. Stirling. 2010. Sea ice: a critical habitat for polar marine mammals and birds, pp 395-424. In D.N. Thomas & G. S. Dieckmann (ed.), Sea Ice, 2nd Edition. Wiley-Blackwell, Chichester, UK.

Varsani, A., S. Kraberger, S. Jennings, E.L. Porzig, L. Julian, M.Massaro, A. Pollard, G. Ballard & Ainley, D.G. 2014. A novel papillomavirus in Adelie penguin (Pygoscelis adeliae) faeces sampled at the Cape Crozier colony, Antarctica. Journal of General Virology 95: 1352–1365 view PDF

Varsani, A., E.L. Porzig, S. Jennings, S. Kraberger, K. Farkas, L. Julian, M. Massaro, G. Ballard & Ainley, D.G. 2015. Identification of an avian polyomavirus associated with Adélie penguins (Pygoscelis adeliae). Journal of General Virology 96: 851–857. view PDF

Wilson, D.J., P. O’B. Lyver, T.C. Greene, A.L. Whitehead, K.M. Dugger, B.J. Karl, J.R. F. Barringer, R. McGarry, A. M. Pollard & Ainley, D.G. 2016. South Polar Skua breeding populations in the Ross Sea assessed from demonstrated relationship with Adélie Penguin numbers. Polar Biology DOI 10.1007/s00300-016-1980-4. view PDF


Foraging Ecology

Ainley, D.G. & G. Ballard. 2011. Non-consumptive factors affecting foraging patterns in Antarctic penguins: a review and synthesis. Polar Biology 35:1–13. view PDF

Ainley, D.G., G. Ballard, R.M. Jones, D. Jongsomjit, S.D. Pierce, W.O. Smith, Jr., S. Veloz. 2015. Trophic cascades in the western Ross Sea, Antarctica: revisited. Marine Ecology Progress Series, doi 10.3354/meps11394. view PDF

Ainley, D.G., G. Ballard, K.J. Barton, B.J. Karl, G.H. Rau, C.A. Ribic & P.R. Wilson. 2003. Spatial and temporal variation of diet composition and quality within a presumed metapopulation of Adélie Penguins. Condor 105: 95-106. view PDF

Ainley, D.G., G. Ballard & K.M. Dugger. 2006. Competition among penguins and cetaceans reveals trophic cascades in the Ross Sea, Antarctica. Ecology, 87:2080–2093. view PDF

Ainley, D.G., G. Ballard, S. Ackley, L.K. Blight, J.T. Eastman, S.D. Emslie, A. Lescroël, S. Olmastroni, S.E. Townsend, C.T. Tynan, P. Wilson, E. Woehler. 2007. Paradigm Lost, or, is top-down forcing no longer significant in the Antarctic Marine Ecosystem? Antarctic Science 19: 283-290. view PDF

Ainley, D.G., C.A. Ribic, G. Ballard, S. Heath, I. Gaffney, B.J. Karl, K.R. Barton, P.R. Wilson & S. Webb. 2004. Geographic structure of Adélie Penguin populations: size, overlap and use of adjacent colony-specific foraging areas. Ecological Monographs 74: 159-178. view PDF

Ainley, D.G., P.R. Wilson, K.R. Barton, G. Ballard, N. Nur & B. Karl. 1998.  Diet and foraging effort of Adélie penguins in relation to pack-ice conditions in the southern Ross Sea.  Polar Biology 20: 311-319. view PDF

Ballance, L.T., D.G. Ainley,  G. Ballard & K. Barton. 2009.  An energetic correlate between colony size and foraging effort in seabirds, an example of the Adélie penguinPygoscelis adeliae. Journal of Avian Biology 40: 279-288. view PDF

Ballard, G., K.M. Dugger, N. Nur & Ainley, D.G. 2010. Foraging strategies of Adélie penguins: adjusting body condition to cope with environmental variability. Marine Ecology Progress Series 405: 287–302. view PDF

Ballerini, T., E.E. Hofmann, D.G. Ainley,  K. Daly, M. Marrari, C.A. Ribic, W.O. Smith Jr., & J.H. Steele. 2013. Productivity and linkages of the food web of the southern region of the western Antarctic Peninsula continental shelf. Progress in Oceanography, view PDF

Ford, R.G., D.G. Ainley,  A. Lescroël, P.O’B. Lyver, V. Toniolo & G. Ballard. 2015. Testing assumptions of central place foraging theory: a study of Adélie penguinsPygoscelis adeliaein the Ross Sea. Journal of Avian Biology, 46: 193-205, doi: 10.1111/jav.00491 view PDF

Jennings, S., A. Varsani, K.M. Dugger, G. Ballard & D.G. Ainley. 2016. Sex-based differences in Adélie Penguin (Pygoscelis adeliae) chick growth rates and diet. PLoS ONE 11(3): e0149090. doi:10.1371/journal.pone.0149090 view PDF

Karnovsky, N., Ainley, D.G. & P. Lee. 2007. The impact and importance of production in polynyas to top-trophic predators: three case histories. In Polynyas (W.O. Smith, Jr. & D.G. Barber, Eds.). Elsevier Publishers, London. Pp 391-410.

Lescroël, A., G. Ballard, V. Toniolo, K.J. Barton, P.R. Wilson, P. O’B. Lyver & D.G. Ainley. 2010. Working less to gain more: when breeding quality relates to foraging efficiency. Ecology 91: 2044–2055 view PDF

Lyver, P.O’B., C.J. MacLeod, G. Ballard, B.J. Karl, J. Adams, D.G. Ainley & P.R. Wilson. 2011. Intra-seasonal variation in foraging behavior among Adélie Penguins (Pygoscelis adeliae) breeding at Cape Hallett, Ross Sea, Antarctica. Polar Biol 34:49–67. view PDF

Sergio, F., O.J. Schmitz, C.J. Krebs, R.D. Holt, M.R. Heithaus, A.J. Wirsing, W.J. Ripple, E. Ritchie, D. Ainley, D. Oro, Y. Jhala, F. Hiraldo and E. Korpimäki. 2014. Towards a cohesive, holistic view of top predation: a definition, synthesis and perspective. Oikos doi: 10.1111/oik.01468 view PDF

Whitehead, A.L., P.O’B. Lyver, G. Ballard, K. Barton, B.J. Karl, K.M. Dugger, S. Jennings, A. Lescroël, P.R. Wilson & D.G. Ainley. 2015. Factors driving Adélie penguin chick size, mass and condition at colonies of differing size in the southern Ross Sea. Marine Ecology Progress Series 523: 199–213. view PDF


Climate Change

Ainley, D.G., G. Ballard, S.D. Emslie, W.R. Fraser, P.R. Wilson & E.J. Woehler. 2003. Adélie Penguins and environmental change. Science 300: 429. view PDF

Ainley, D.G., E.D. Clarke, K. Arrigo, W.R. Fraser, A. Kato, K.J. Barton & P.R. Wilson. 2005. Decadal-scale changes in the climate and biota of the Pacific sector of the Southern Ocean, 1950s to the 1990s. Antarctic Science 17: 171-182. view PDF

Ainley, D.G., K.A. Hobson, X. Crosta, G.H. Rau, L.I. Wassenaar & P.C. Augustinus. 2006. Holocene variation in the Antarctic coastal food web: linking δD and δ13Cin snow petrel diet and marine sediments. Marine Ecology Progress Series, 306: 31-40. view PDF

Ainley, D.G., J. Russell, S. Jenouvrier, E. Woehler, P. O’b. Lyver, W.R. Fraser, G.L. Kooyman. 2010. Antarctic penguin response to habitat change as earth’s troposphere reaches 2°c above pre-industrial levels. Ecological Monographs 80: 49-66. view PDF

Ainley, D.G., P.R. Wilson & W.R. Fraser. 2001. Effects of climate change on Antarctic sea ice and penguins. Pp. 26-27inImpacts of climate change on wildlife (R.E. Green, M. Harley, M. Spalding & C. Zockler, eds.). RSPB, Cambridge, UK.

Arrigo, K.R., G.L. van Dijken, D.G. Ainley, M.A. Fahnestock & T. Markus. 2002. The impact of the B-15 iceberg on productivity and penguin breeding success in the Ross Sea, Antarctica. Geophys. Res. Letters 29(7), 10.1029/2001GLO14160. view PDF

Ballard G., V. Toniolo, D.G. Ainley, C.L. Parkinson, K.R. Arrigo & P.N. Trathan. 2010. Responding to climate change: Adélie penguins confront astronomical and ocean boundaries. Ecology 91: 2056-2069. view PDF

Dugger, K.M., G. Ballard, D.G. Ainley, P.O’B. Lyver & C. Schine. 2014. Adélie penguins coping with environmental change: results from a natural experiment at the edge of their breeding range. Frontiers in Ecology and Evolution 2:68, doi: 10.3389/fevo.2014.00068. view PDF

Emslie, S.D., P.A. Berkman, D.G. Ainley, L. Coats & M. Polito. 2003. Late-Holocene initiation of ice-free ecosystems in the southern Ross Sea, Antarctica. Marine Ecology Progress Series 262: 19-25. view PDF

Kooyman, G.L., D.G. Ainley,  G. Ballard and P.J. Ponganis. 2007. Effects of giant icebergs on two emperor penguin colonies in the Ross Sea, Antarctica. Antarctic Science 19: 31-38. view PDF

LaRue, M.A., D.G. Ainley, M. Swanson, K.M. Dugger, P.O. Lyver, K. Barton & G. Ballard. 2013. Climate change winners: receding ice fields facilitate colony expansion and altered dynamics in an Adélie Penguin metapopulation. PLoS ONE 8(4): e60568. doi:10.1371/journal.pone.0060568. view PDF

Lescroël, A., G. Ballard, D. Grémillet, M. Authier & Ainley, D.G. 2014. Antarctic climate change: extreme events disrupt plastic response in Adélie penguins. PLoS ONE, 9(1): e85291. doi:10.1371/journal.pone.0085291. view PDF

Shepherd, L.D., C.D. Millar, G. Ballard, D.G. Ainley,  P.R. Wilson, G.D. Haynes, C. Baroni & D.M. Lambert. 2005. Microevolution and mega-icebergs in the Antarctic. Proc. Natl Acad. Sci. 102 (46): 16717-16722. view PDF

Smith, R.C., E. Domack, S. Emslie, W.R. Fraser, D.G. Ainley,  K. Baker, J. Kennett, A. Leventer, E. Mosley-Thompson, S. Stammerjohn & M. Vernet. 1999. Marine ecosystem sensitivity to historical climate change: Antarctic Peninsula.  BioScience 49: 393-404.

Wilson, P.R., D.G. Ainley,  N. Nur, S.S. Jacobs, K.J. Barton, G. Ballard & J.C. Comiso. 2001. Adélie Penguin population change in the Pacific Sector of Antarctica: Relation to sea-ice extent and the Antarctic Circumpolar Current. Marine Ecology Progress Series 213: 301-309. view PDF


Ross Sea Ecology

Ainley, D.G. 2002. The Ross Sea, Antarctica: Where all ecosystem processes still remain for study. CCAMLR Document WG-EMM 02/60. Hobart, Australia. view PDF

Ainley, D.G. 2002. The Ross Sea, Antarctica: Where all ecosystem processes still remain for study, but maybe not for long. Marine Ornithology 31: 55-62. view PDF

Ainley, D.G. 2004. Acquiring a ‘base datum of normality’ for a marine ecosystem: the Ross Sea, Antarctica. CCAMLR document WG-EMM-04/20, Hobart. view PDF

Ainley D.G. 2007. Insights from study of the last intact neritic marine ecosystem. Trends in Ecology & Evolution 22: 444-445. view PDF

Ainley, D.G. 2010. A history of the exploitation of the Ross Sea, Antarctica. Polar Record46 (238): 233–243. view PDF

Ainley, D., V. Toniolo, G. Ballard, K. Barton, J. Eastman, B. Karl, S. Focardi, G. Kooyman, P. Lyver, S. Olmastroni, B.S. Stewart, J.W. Testa & P. Wilson. 2006. Managing Ecosystem Uncertainty: Critical Habitat and Dietary Overlap of Top-Predators in the Ross Sea. CCAMLR document EMM 06 – 07, Hobart. view PDF

Ainley, D.G., G. Ballard & J. Weller. 2010. ROSS SEA BIOREGIONALIZATION Part I: Validation of the 2007 CCAMLR Bioregionalization Workshop Results Towards Including the Ross Sea in a Representative Network of Marine Protected Areas in the Southern Ocean. CCAMLR WG-EMM-10/11, Hobart. view PDF

Ballard, G., D. Jongsomjit & D.G. Ainley. 2010. ROSS SEA BIOREGIONALIZATION, Part Ii: Patterns of Co-Occurrence of Mesopredators in an Intact Polar Ocean Ecosystem. CCAMLR WG-EMM-10/12, Hobart. view PDF

Ballard, G., Jongsomjit, D., Veloz, S.D., Ainley, D.G. 2012. Coexistence of mesopredators in an intact polar ocean ecosystem: The basis for defining a Ross Sea marine protected area. Biological Conservation 156: 72-82. view PDF

Smith, W.O. Jr., D.G. Ainley & R. Cattaneo-Vietti. 2006. Marine Ecosystems: the Ross Sea.InAntarctic Ecology: From Genes to Ecosystems, Phil. Trans. R. Soc. B (2007) 362, 95–111. view PDF

Smith, W.O. Jr., D.G. Ainley, R. Cattaneo-Vietti & E.E. Hofmann. 2012. The Ross Sea continental shelf: Regional biogeochemical cycles, trophic interactions, and potential future changes. In: Antarctic Ecosystems: An Extreme Environment in a Changing World (A.D. Rogers, N.M. Johnston, E.J. Murphy and A. Clarke, Eds.). Chapter 7, Blackwell Publishing, London, pp. 213-242. view PDF

Smith, W.O. Jr., D.G. Ainley,  K.R. Arrigo & M.S. Dinniman. 2014. The oceanography and ecology of the Ross Sea. Annual Review of Marine Science 6:469–487. view PDF


Whales (Penguin competitor)

Ainley, D.G., K.M. Dugger, V. Toniolo & I. Gaffney. 2007. Cetacean occurrence patterns in the Amundsen and southern Bellingshausen Sea sector, Southern Ocean. Marine Mammal Science 23: 287-305. view PDF

Ainley, D.G., G. Ballard, S. Olmastroni. 2009. An Apparent Decrease in the Prevalence of “Ross Sea Killer Whales” in the Southern Ross Sea. Aquatic Mammals 35(3), 335-347. view PDF

Ainley, D.G., G. Ballard, L.K. Blight, S. Ackley, S.D. Emslie, A. Lescroël, S. Olmastroni, S.E. Townsend, C.T. Tynan, P. Wilson, E. Woehler. 2010. Impacts of cetaceans on the structure of southern ocean food webs. Marine Mammal Science 26: 482-489. view PDF

Ainley, D.G., D. Jongsomjit, G. Ballard, D. Thiele, W.R. Fraser, C.T. Tynan. 2012. Modeling the relationship of Antarctic minke whales to major ocean boundaries. Polar Biology 35:281-290. view PDF

Ainley, D.G. & G. Ballard. 2012. Trophic Interactions and the decrease in Killer Whale (Orcinus orca) prevalence with reduced availability of large fish in the southern Ross Sea. Aquatic Mammals 38: 153-160. view PDF

Ainley, D.G., K. Lindke, G. Ballard, P.O’B. Lyver, S. Jennings, V. Toniolo, J. Pennycook, A. Lescroël, M. Massaro & J.A. Santora. Spatio-temporal occurrence patterns of cetaceans near Ross Island, Antarctica, 2002-2015: implications for foodweb dynamics. Polar Biology, submitted

Ballard, G. & D.G. Ainley. 2005. Killer whale harassment of Adélie penguins at Ross Island. Antarctic Science17: 385-386 view PDF

Lauriano, G., M. Vacchi, D. Ainley & G. Ballard. 2007. Observations of top predators foraging on fish in the pack ice of the southern Ross Sea. Ant. Sci. 19: 439-440. view PDF


Seals (Penguin competitor, Predator)

Ainley, D.G., G. Ballard, B.J. Karl & K.T. Dugger. 2005. Leopard seal predation rates at penguin colonies of different size. Antarctic Science17: 335-340. view PDF

Ainley, D. G., M.A. LaRue, I. Stirling, S. Stammerjohn & D.B. Siniff. 2015. An apparent population decrease, or change in distribution, of Weddell seals along the Victoria Land coast. Marine Mammal Science, DOI: 10.1111/mms.12220. view PDF

Ainley, D.G. & D.B. Siniff. 2009. The importance of Antarctic toothfish as prey of Weddell Seals in the Ross Sea: A Review. Antarctic Science 21: 317-327. view PDF

Kim, S.Z., D.G. Ainley,  J. Pennycook & J.T. Eastman. 2011. Antarctic toothfish heads found along tide cracks of the McMurdo Ice Shelf. Antarctic Science 23: 469-470. view PDF

LaRue, M.A., J.J. Rotella, R.A. Garrott, D.B. Siniff,  D.G. Ainley,  G.E. Stauffer, C.C. Porter & P.J. Morin. 2011. Satellite imagery can be used to detect variation in abundance of Weddell seals (Leptonychotes weddellii) in Erebus Bay, Antarctica. Polar Biology 34:1727-1737. view PDF

Salas, L., N. Nur, D. Ainley, J. Burns, J. Rotella, G. Ballard. 2016. Coping with loss of large, energy-dense prey: A potential bottleneck for Weddell Seals in the Ross Sea. Ecological Applications, in press. view PDF

Siniff, D.B., R.A. Garrott, J.J. Rotella, W.R. Fraser & D.G. Ainley. 2008. Projecting the effects of environmental change on Antarctic seals. Antarctic Science 20: 425-435. view PDF


Antarctic Toothfish (penguin competitor)

Abrams, P.A., D.G. Ainley,  L.K. Blight, P.K. Dayton, J.T. Eastman, J.L. Jacquet. 2016. Necessary elements of precautionary management: implications for the Antarctic toothfish. Fish & Fisheries 17, 1152–1174, DOI: 10.1111/faf.12162. view PDF

Ainley, D.G., G. Ballard, J.T. Eastman, C.W. Evans, N. Nur, C.L. Parkinson. 2016. Changed prevalence, not absence explains toothfish status in McMurdo Sound. Antarctic Science, doi:10.1017/S0954102016000584 view PDF

Ainley, D.G., N. Nur, J.T. Eastman, G. Ballard, C.L. Parkinson, C.W. Evans, A.L. DeVries. 2013 Decadal trends in abundance, size and condition of Antarctic toothfish in McMurdo Sound, Antarctica, 1972-2011. Fish & Fisheries 14, 343–363 DOI: 10.1111/j.1467-2979.2012.00474.x. view PDF

Ainley, D.G., C.M. Brooks, J.T. Eastman, and M. Massaro. 2012. Unnatural selection of Antarctic Toothfish in the Ross Sea, Antarctica. F. Huettmann (ed.),Protection of the Three Poles, pp. 53-75. DOI 10.1007/978-4-431-54006-9_3, Springer, Tokyo. view PDF

Ainley, D.G., J.T. Eastman, C.M. Brooks. 2015. Comments on ‘‘The Antarctic toothfish (Dissostichus mawsoni): biology, ecology, and life history in the Ross Sea region,’’ by S. Hanchet et al. Hydrobiologia, 2015, 761:397–414. view PDF


Human Impacts

Ainley, D.G. & L.K. Blight. 2009. Ecological repercussions of historical fish extraction from the southern ocean. Fish and Fisheries 10: 13-38.Blight, L.K, and Ainley, D.G. 2008. Southern Ocean not so pristine. Science 321: 1443. view PDF

Ainley, D.G. & C.M. Brooks. 2013. Exploiting the Southern Ocean: rational use or return to tragedy of the commons, pp. 143-154.InLiggett, D. & Hemmings, A. (ed.) Exploring Antarctic Values. Antarctic Gateway Publication Series, No. 1031. Christchurch NZ. view PDF

Ainley, D.G. & D. Pauly. 2014. Fishing down the food web of the Antarctic continental shelf and slope. Polar Record 50: 92-107,doi:10.1017/S0032247412000757. view PDF

Ainley, D.G. & D. Pauly. 2016. Antarctica. In: Global Atlas of Marine Fisheries: A Critical Appraisal of Catches and Ecosystem Impacts (D. Pauly & D. Zeller, Eds.). Island Press, Washington, DC. P. 188.

Blight, L.K., D.G. Ainley,  S.F. Ackley, G. Ballard, T. Ballerini, R.L. Brownell Jr., C.-H. C. Cheng, M. Chiantore, D. Costa, M.C. Coulter, P. Dayton, A.L. Devries, R. Dunbar, S. Earle, J.T. Eastman, S. D. Emslie, C.W. Evans, R.A. Garrott, S. Kim, G. Kooyman, A. Lescroël, M. Lizotte, M. Massaro, S. Olmastroni, P.J. Ponganis, J. Russell, D.B. Siniff, W.O. Smith Jr., B.S. Stewart, I. Stirling, J. Willis, P.R Wilson & E.J. Woehler. 2010. Fishing for data in the Ross Sea. Science 330: 1316. view PDF

Brooks, C.M. & D.G. Ainley. 2016. Fishing the bottom of the earth: the political challenges of ecosystem-based management. In: Handbook on the Politics of Antarctica (K. Dodds, A.D. Hemmings, P. Roberts, Eds.), in press.

Brooks, C.M., L.B. Crowder, L.M. Curran, R.B. Dunbar, D.G. Ainley, K.J. Dodds, K.M. Gjerde & U.R. Sumaila. 2016. The decline of science-based management in the Southern Ocean: Conservation or compromise? Science 354: 185-187. view PDF

Christian, C., D. Ainley, M. Bailey, P. Dayton, J. Hocevar, M. LeVine, J. Nikoloyuk, C. Nouvian, E. Velarde, R. Werner & J. Jacquet. 2013. A review of formal objections to Marine Stewardship Council fisheries certifications. Biological Conservation 161: 10–17. view PDF

Christian, C., D. Ainley, M. Bailey, P. Dayton, J. Hocevar, M. LeVine, J. Nikoloyuk, C. Nouvian, E. Velarde, R. Werner & J. Jacquet. 2013. Not ‘‘the best environmental choice in seafood’’: A response to Gutiérrez and Agnew (2013). Biological Conservation 161: view PDF

Jacquet, J., D. Pauly, D. Ainley, S. Holt, P. Dayton & J. Jackson. 2010. Seafood stewardship in crisis. Nature 467: 28-29. view PDF

Jacquet, J., E. Blood-Patterson, C. Brooks & D. Ainley. 2016. ‘Rational use’ in Antarctic waters. Marine Policy 63: 28-34. view PDF

Marschoff, E.R., E.R. Barrera-Oro, N.S. Alescio & D.G. Ainley. 2012. Slow recovery of previously depleted demersal fish at the South Shetland Islands, 1983–2010. Fisheries Research 125/126: 206-213. view PDF

Tin, T., Z.L. Fleming, K.A. Hughes, D.G. Ainley, P. Convey, C.A. Moreno, S. Pfeiffer, J. Scott & I. Snape. 2009. Impacts of local human activities on the Antarctic environment. Antarctic Science 21, doi:10.1017/S0954102009001722 view PDF

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